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A former CIA agent who participated in the UA virus attack and now owns a cosplay cafe in Shanghai taking care of Hakkoh
Santana thought he was doing the right thing when he worked with the CIA, injecting the residents of a small Chinese village with what he assumed was an anti-viral drug to protect the people from the Ua Virus. However, he found out that he was simply another pawn in America's game. In reality, Santana and his fellow Americans were injecting the villagers with a drug that would cause the Ua Virus to mutate their bodies. Several people died, and those that survived were taken to The Factory run by the Snakes to be experimented on further, where they would later become Borners or Unblooms.
Having realized his part in this scheme, Santana went AWOL from the CIA and from his friends, and stuck around in China. However, not too long afterwards, he found an escaping woman and had an attack of conscience. When the Snakes caught up with her, he returned with them to the Factory and begged Alphard herself for Hakko's release. Surprisingly, Alphard let the two of them go, while Liang Qi taunted Santana that Hakkoh would never be able to speak or cry out in pleasure with her mutation. Once he was out of their grasp, he simply tried to hide in Shanghai by setting up a Japanese-style cosplay and karaoke bar and taking care of Hakkoh, though he suspected that she still blamed him for his part in the conspiracy.
Even so, events conspired to bring both himself and Hakkoh back into trouble. First when the Unblooms were wandering around Shanghai during a festival, then again when Minoru caught the mark of the Borner next to Hakkoh's breast. Eventually, he was convinced by a fellow spy to help out Canaan as she traveled with Maria and Minoru to the village where his troubles began. He traveled with the group to the Factory, hoping to expose everything the CIA had done there and reveal it to the public. However, he was captured by Liang Qi and then accidentally killed by Hakkoh. Before his mind was completely gone, he begged Hakkoh to tell him "I love you," which she did out loud until his brain exploded.