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Twin princesses Fine and Rein of the Sunny Kingdom are said to be the "most unprincesslike princesses ever known in Mystery Star". Grace, the person who saved Mystery Star once, chose them to save Mystery Star again. She gave them the Prominence, which levels up with them as they get more and more princesslike. They meet many people and have Princess Parties, in which they try to win "Best Princess" titles. They usually end up messing up, though, in the Parties. They meet Eclipse, who they think is evil, but is he really? They also meet Bright, who has a crush on Fine. Rein has a crush on Bright, though. They meet princesses and princes of the other seven countries later. But are there some unknown enemies for Fine and Rein?